95 –98. Bacterial Diversity in Subbituminous Coal and Soil from Coal Mine of South Sumatra, Indonesia

International Journal of Integrative Biology IJIB, 2013, Vol. 14, No. 2, 102 ©OmicsVista Group, All rights reserved derived fro m bacteria trapped during sedimentation Bo ivin-Jahns et al., 1996. It suggests that the extraction method, particu larly the direct method Zhou et al. , 1996, is more suitable to be applied on the high clay soil. It successfully obtained OTUs that might not be obtained by culture method. Firmicutes, especially Bacillus, has higher abundance of OTU than others in their co mmun ity in the coal sample. It is possible since the genus Bacillus is an r- strategist organism Sarathchandra et al., 1997. In rich environments of organic matter such as coal, Bacillus easily outperforms other members by intensifying its growth by using abundant organic resources. The high abundance of Bacillus showed that they had been in the favorable physico-chemical conditions in the coal. Moreover, they are well known to have an endospore as defense mechanism of sustainability. In conclusion, the differences in physical and chemical properties of the coal and the surrounding soil bacterial showed different diversity in their co mmunit ies though they still have genetic relatedness. Conflict of interest The authors declare no conflict of interest. Acknowledgement This research was hardly carried out without support from PhD Scholarship Grant Number: BPPS Dikti-ITB 2009 Beasiswa Pendidikan Pasca Sarjana, Directorate General of Higher Education, M inistry of National Education, Indonesia, and Dissertation Support from State Islamic University Syarif Hidayatullah Jakarta, M inistry of Religious Affairs, Indonesia to M RP. We also appreciated to Ir. Eko Pujiantoro and his staff from PT. Bukit Asam Company.. References Aminin ALN, Warganegara FM, Aditiawati P and Akhmaloka 2008a Culture-independent and culture-dependent approaches on microbial community analysis at Gedongsongo analysis at Gedongsongo GS-2 hot spring. Int. J. Integrative Biol., 2: 145-152. Aminin ALN, Warganegara FM, Aditiawati P and Akhmaloka 2008b Simple enrichment and independent cultures to expand bacterial community analysis from Gedongsongo hot spring. J. Biosci. Bioeng ., 106: 211-214. Andreetta A, Macci C, Ceccherini MT, Cecchini G, Masciandaro G, Pietramellara G and Carnicelli S 2012 Microbial dynamics in Mediterranean moder humus. Biology and Fertility of Soils, 48: 259- 270. Beckmann S, Lueders T, et al. 2011 Acetogens and acetoclastic methanosarcinales govern methane formation in abandoned coal mines. Appl. Environ. Microbiol., 7711: 3749-3756. Berga M, Sze´kely AJ, Langenheder S 2012 Effects of disturbance intensity and frequency on bacterial community composition and function. PLoSONE 7: e36959. doi:10.1371 journal.pone.0036959 Boivin-Jahns V, Ruimy R, Bianchi A, Daumas S and Christen R 1996 Bacterial diversity in a deep-subsurface clay environment. Appl. Environ. Microbiol ., 62: 3405-3412. Carrigg C, Rice O, Kavanagh S, Collins G, and O’Flaherty V 2007 DNA extraction method affects microbial community profiles from soils and sediment. Appl. Microbial. and Biotechnol., 77: 955-964. Daniel R 2005 The metagenomics of soil. Nat. Rev.Microbiol. 3: 470-478. Davison J, Öpik M, Zobel M, Vasar M, Metsis M, et al 2012 Communities of arbuscular mycorrhizal fungi detected in forest soil are spatially heterogeneous but do not vary throughout the growing season. PLoSONE 7: e41938. doi:10.1371journal.pone.0041938 Dhand NK, Toribio JAL and Whittington RJ 2009 Adsorption of Mycobacterium avium subsp. paratuberculosis to soil particles. Appl. Environ. Microbiol ., 75: 5581-5585. Dojka MA, Harris JK and Pace NR 2000 Expanding the known diversity and environmental distribution of an uncultured phylogenetic division of bacteria. Appl. Environ. Microbiol., 66: 1617-1621. Feinstein LM, Sul WJ and Blackwood CB 2009 Assessment of bias associated with incomplete extraction of microbial DNA from soil. Appl. Environ. Microbiol ., 75: 5428-5433. Felske A, Engelen B, Nübel U and Backhaus H 1996 Direct ribosome isolation from soil to extract bacterial rRNA for community analysis. Appl. Environ. Microbiol., 62: 4162-4167. Ferrenberg S, ONeill SP, Knelman JE, Todd B, Duggan S, Bradley D, ... and Nemergut DR 2013 Changes in assembly processes in soil bacterial communities following a wildfire disturbance. The ISME journal. Fromin N, Hamelin J, Tarnawski S, Roesti D, Jourdain ‐Miserez K, Forestier N, T eyssier ‐Cuvelle S, Gillet F, Aragno M and Rossi P 2002 Statistical analysis of denaturing gel electrophoresis DGE fingerprinting patterns. Environmental Microbiology 4: 634-643. Ge Y, Chen C, Xu Z, Oren R and He JZ 2010 The spatial factor, rather than elevated CO2, controls the soil bacterial community in a temperate forest ecosystem. Appl. Environ. Microbiol., 76: 7429- 7436. Ghose MK 2004 Effect of opencast mining on soil fertility. J. Sci. Ind. Res. , 63: 1006-1009. Gonzalez A, Clemente JC, et al. 2011 Our microbial selves: what ecology can teach us. EMBO Reports, 12: 775-784. Haering KC, Daniels WL and Galbraith JM 2004 Appalachian mine soil morphology and properties. Soil Sci. Soc. Am. J., 684: 1315- 1325. Hall A 1999 Bioedit: a user-friendly biological sequence alignment editor and analysis program for Windows 9598 NT . Nucl. Acids. Sym p. Ser.

41: 95 –98.

Harris JK, Kelley ST and Pace NR 2004 New perspective on uncultured bacterial phylogenetic division OP11. Appl. Environ. Microbiol. , 702: 845-849. Hong Z, Rong X, Cai P, Dai K, Liang W, Chen W and Huang Q 2012 Initial adhesion of Bacillus subtilis on soil minerals as related International Journal of Integrative Biology IJIB, 2013, Vol. 14, No. 2, 103 ©OmicsVista Group, All rights reserved to their surface properties. European Journal of Soil Science, 63: 457-466. İnceoǧlu Ö, Hoogwout EF, Hill P and van Elsas JD 2010 Effect of DNA extraction method on the apparent microbial diversity of soil. Appl. Environ. Microbiol ., 76: 3378-3382. Jiang D, Huang Q, Cai P, Rong X and Chen W 2007 Adsorption of Pseudomonas putida on clay minerals and iron oxide. Colloids and Surfaces B: Biointerfaces , 54: 217-221. Konstantinov SR, Zhu WY, Williams BA, Tamminga S, Vos WM and Akkermans AD 2003 Effect of fermentable carbohydrates on piglet faecal bacterial communities as revealed by denaturing gradient gel electrophoresis analysis of 16S ribosomal DNA. FEMS Microbiology Ecology , 43: 225-235. Laplante K and Derome N 2011 Parallel changes in the taxonomical structure of bacterial communities exposed to a similar environmental disturbance. Ecology and Evolution 1: 489 –501. Limberger R and Wickham SA 2012 Disturbance and diversity at two spatial scales. Oecologia 168: 785 –795. DOI 10.1007s00442- 011-2140-8. Machulla G, Bruns MA and Scow KM 2005 Microbial properties of mine spoil materials in the initial stages of soil development. Soil Sci. Soc. Am . J. , 694: 1069-1077. Morales SE and Holben WE 2009 Empirical testing of 16S rRNA gene PCR primer pairs reveals variance in target specificity and efficacy not suggested by in silico analysis. Appl. Environ. Microbiol. 75 : 2677-2683. Muirhead RW, Collins RP and Bremer PJ 2006 Interaction of Escherichia coli and soil particles in runoff. Appl. Environ. Microbiol ., 72: 3406-3411. Nakatsu CH, Torsvik V and Øvreås L 2000 Soil community analysis using DGGE of 16S rDNA polymerase chain reaction products. Soil Science Society of America Journal, 644: 1382-1388. Norris T B, Wraith JM, Castenholz RW and McDermott TR 2002 Soil microbial community structure across a thermal gradient following a geothermal heating event. Appl. Environ. Microbiol., 68: 6300-6309. Rappé MS and Giovannoni SJ 2003 The uncultured microbial majority. Annual Reviews in Microbiology 57: 369-394. Rodrigue JA and Burger JA 2004 Forest soil productivity of mined land in the midwestern and eastern coalfield regions. Soil Sci. Soc. Am . J. , 683: 833-844. Rumpel C and Kögel-Knabner I 2004 Microbial use of lignite compared to recent plant litter as substrates in reclaimed coal mine soils. Soil Biol. Biochem., 361: 67-75. Salam LB, Obayori OS, Akashoro OS and Okogie GO 2011 Biodegradation of bonny light crude oil by bacteria isolated from contaminated soil. Int. J. Agric. Biol., 13: 245 –250. Santos HF, Carmo FL, Leite DCA, Jesus HE, De Carvalho Maalouf P, Almeida C, Soriano A. U. et al. 2012 Comparison of different protocols for the extraction of microbial DNA from reef corals. Brazilian Journal of Microbiology

43: 517-527.