Species Of The Genus Calamus (Palmae) Section Phyllanthectus Furtado In Sumatera
SPECIES OF THE GENUS CALAMUS (PALMAE) SECTION
PHYLLANTHECTUSFURTADO IN SUMATRA
IDRAMSA
NRP.97268
POST GRADUATE PROGRAM
BOGOR INSTITUTE OF AGRICULTURE
2000
11
ABSTRACT
The taxonomy of Calamus section Phyllanthectus in Sumatra was studied
based on morphological observation on 57 specimens deposited in the Herbarium
Bogoriense. The section consists of C. asperrimus, C. axillaris, C. caesius, C.
laevigalus, C. manan, C. oxleyanus var. oxleyanus, C. oxleyanus var. obovalus,
C. palel71banicus, C. tumidus, C. uni(arius var. uni(arius and C. uni(arius var.
siberulensis. The latter is proposed here for the first.
III
ABSTRAK
Taksonomi
Calamus
seksi
Phyllanthectus di
Sumatera dipelajari
berdasarkan pengamatan morfologi terhadap 57 nomor koleksi (terdiri atas 209
lembar) yang terdapat di Herbarium Bogoriense. Di Sumatera seksi ini terdiri atas
C. asperrimus, C. axillaris, C. caesius, C. laevigatus, C. manan, C. ox/eyanus var.
ox/eyanus, C. ox/eyanus var. obovatus, C. palembanicus, C. tumidus, C. uni(arius
var. 1I11i(arius dan C. unifarius var. siberutensis. Varietas yang disebutkan terakhir
diusulkan pertama sekali sebagai takson baru.
IV
SPECIES OF THE GENUS CALAMUS (PALMAE) SECTION
PHYLLANTHECTUSFURTADO IN SUMATRA
IDRAMSA
NRP. 97268
A Thesis Submitted for the Degree of Master
At
Post Graduate Program Bogor Institute of Agriculture
POST GRADUATE PROGRAM
BOGOR INSTITUTE OF AGRICULTURE
2000
v
Thesis Title
Name
NRP.
Study Program
: SPECIES OF THE GENUS CALAMUC; (PALMAE)
SECTION PHYLLANT11ECT[fS FURTADO IN
SUMATRA
:ldramsa
: 97268
: Biology
Approved by
1. Supervisor Committee
(Prof. Dr. Ir. Edi Guhardia. M.Sc.)
Supervisor
(Prof. Dr. Mien A. Rifail
Supervisor
2. Chair Person
Study Program
(Dr. II'. Dede Seliadi, MS)
Day of graduation: October 21. 2000
ＭＺ＿ＦｾＬ､｜ｊ｡ｴ｣＠
(Dr. Johanis P. Mogea)
Supervisor
Program
vi
CURRICULUM VITAE
Idramsa is a son ofPandi bin H. Idris and Fatimah. He was born in Medan
on 20 September 1971.
In 1985 he was graduated from Elementary School, in 1988 from Junior
High School and in 1991 from Senior High School all in Labuhan Deli. In 1996
he was graduated from the Faculty of Education and Natural Science !KIP Medan
majoring in Biology.
in September, 1997 he registered as a student at the Graduate Program IPB
Bogor, studying Plant Taxonomy.
Vll
ACKNOWLEDGEMENTS
Many people contributed in this finishing study. As I finished writing this
paper, I am very grateful to all of them. Some people and institutions have been
particularly helpful and I would like to mention them individually.
Firstly I would like to thank my supervisors Prof. Dr. Ir. Edi Guhardja,
M.Sc., Prof. Dr. Mien A. Rifai and Dr. Johanis P. Mogea who give me constant
support, advice, and encouragement during my study.
I would like to express my gratitude as well to the Herbarium Bogoriense
(LIPI), for granting me pennisson to conduct research and providing some
facilities. I wish to thank the following person, Dr. Arie Budiman, the Head
Puslitbang Biologi; Dr. Irawati, Head of Herbariwn Bogoriense; Dr. Dedy
Darnaedi, Head of Bogor Botanical Garden; Prof. Dr. Ir. Syafrida Manuwoto,
M. Sc., Director of the Graduate School Bogor Institute of Agriculture; Dra.
Sunarti, and Drs. Uway Warsita Mahyar curator in Herbarium Bogoriense, the
acknowledgement is also go to Dra. Enok, Head of Library in the Herbarium
Bogoriense.
My gratitude is due to Dr. John Dransfield of the Royal Botanic Garden,
Kew for sharing his knowledge and for introducing me to the wonder world of
palms.
I would like to thank to Dr. Harry Wiriadinata, Titik Rugayah, Ir. Himmah
Rustiami, M.Sc. and Teguh Triono Sp. M,Sc., for helping me in various ways.
viii
'1 would like to thank Kartika Indah, S.Pd, M.Si, Dra. Kasrina, If. Nurhaida
Iriani Sinaga, Medi Hendra, S.Si., Andam Surianty Ardan, S.Si., for their moral
supports.
My very special thanks are due to my uncle and aunt (Drs. H. Arifin Saleh
and Dra. Hj. Nurlina Arifin) for giving me continuous moral support and attention
during my study in Bogor, and also to my sister Dra. Riwayati, M.Si, for her
attention and advice.
IX
TABLE OF CONTENTS
ABSTRACT ................................................................................... .ii
CURICULUM VITAE ...................................................................... vi
ACKNOWLEDGEMENTS ................................................................. vii
TABLE OF CONTENTS ................................................................... .ix
LIST OF FIGURES .......................................................................... xi
INTRODUCTION .............................................................................. 1
METHOD OF STUDY .................................................................... 3
RESULT AND DISCUSSION .......................................................... 5
Section P hyllanthectus Furtado ................................................... 7
Key to species of the Genus Calamus (Palmae) section Phyllanrhectus
Furtado in Sumatra .............................................................. 8
Species Description ................................................................. .12
Calamus asperrimlls Blume ............................................. 12
Calamus axillaris Becc ................................................... 13
Calamus caesius Blume .................................................. .15
Calams laevigatus Mart .................................................. 18
Calamus manan Miq ...................................................... 20
Calamus oxleyanus Teysm. et Binn ................................... 22
Calamus palembanicus Becc ........................................... 25
Calamus tllmidus Furtado ............................................... 26
Calamus unifarius H. Wend!.. ......................................... 27
x
CONCLUSION ... ........... ' ............................................................ .34
REFERENCES ............................................. '" ........................... 35
PHYLLANTHECTUSFURTADO IN SUMATRA
IDRAMSA
NRP.97268
POST GRADUATE PROGRAM
BOGOR INSTITUTE OF AGRICULTURE
2000
11
ABSTRACT
The taxonomy of Calamus section Phyllanthectus in Sumatra was studied
based on morphological observation on 57 specimens deposited in the Herbarium
Bogoriense. The section consists of C. asperrimus, C. axillaris, C. caesius, C.
laevigalus, C. manan, C. oxleyanus var. oxleyanus, C. oxleyanus var. obovalus,
C. palel71banicus, C. tumidus, C. uni(arius var. uni(arius and C. uni(arius var.
siberulensis. The latter is proposed here for the first.
III
ABSTRAK
Taksonomi
Calamus
seksi
Phyllanthectus di
Sumatera dipelajari
berdasarkan pengamatan morfologi terhadap 57 nomor koleksi (terdiri atas 209
lembar) yang terdapat di Herbarium Bogoriense. Di Sumatera seksi ini terdiri atas
C. asperrimus, C. axillaris, C. caesius, C. laevigatus, C. manan, C. ox/eyanus var.
ox/eyanus, C. ox/eyanus var. obovatus, C. palembanicus, C. tumidus, C. uni(arius
var. 1I11i(arius dan C. unifarius var. siberutensis. Varietas yang disebutkan terakhir
diusulkan pertama sekali sebagai takson baru.
IV
SPECIES OF THE GENUS CALAMUS (PALMAE) SECTION
PHYLLANTHECTUSFURTADO IN SUMATRA
IDRAMSA
NRP. 97268
A Thesis Submitted for the Degree of Master
At
Post Graduate Program Bogor Institute of Agriculture
POST GRADUATE PROGRAM
BOGOR INSTITUTE OF AGRICULTURE
2000
v
Thesis Title
Name
NRP.
Study Program
: SPECIES OF THE GENUS CALAMUC; (PALMAE)
SECTION PHYLLANT11ECT[fS FURTADO IN
SUMATRA
:ldramsa
: 97268
: Biology
Approved by
1. Supervisor Committee
(Prof. Dr. Ir. Edi Guhardia. M.Sc.)
Supervisor
(Prof. Dr. Mien A. Rifail
Supervisor
2. Chair Person
Study Program
(Dr. II'. Dede Seliadi, MS)
Day of graduation: October 21. 2000
ＭＺ＿ＦｾＬ､｜ｊ｡ｴ｣＠
(Dr. Johanis P. Mogea)
Supervisor
Program
vi
CURRICULUM VITAE
Idramsa is a son ofPandi bin H. Idris and Fatimah. He was born in Medan
on 20 September 1971.
In 1985 he was graduated from Elementary School, in 1988 from Junior
High School and in 1991 from Senior High School all in Labuhan Deli. In 1996
he was graduated from the Faculty of Education and Natural Science !KIP Medan
majoring in Biology.
in September, 1997 he registered as a student at the Graduate Program IPB
Bogor, studying Plant Taxonomy.
Vll
ACKNOWLEDGEMENTS
Many people contributed in this finishing study. As I finished writing this
paper, I am very grateful to all of them. Some people and institutions have been
particularly helpful and I would like to mention them individually.
Firstly I would like to thank my supervisors Prof. Dr. Ir. Edi Guhardja,
M.Sc., Prof. Dr. Mien A. Rifai and Dr. Johanis P. Mogea who give me constant
support, advice, and encouragement during my study.
I would like to express my gratitude as well to the Herbarium Bogoriense
(LIPI), for granting me pennisson to conduct research and providing some
facilities. I wish to thank the following person, Dr. Arie Budiman, the Head
Puslitbang Biologi; Dr. Irawati, Head of Herbariwn Bogoriense; Dr. Dedy
Darnaedi, Head of Bogor Botanical Garden; Prof. Dr. Ir. Syafrida Manuwoto,
M. Sc., Director of the Graduate School Bogor Institute of Agriculture; Dra.
Sunarti, and Drs. Uway Warsita Mahyar curator in Herbarium Bogoriense, the
acknowledgement is also go to Dra. Enok, Head of Library in the Herbarium
Bogoriense.
My gratitude is due to Dr. John Dransfield of the Royal Botanic Garden,
Kew for sharing his knowledge and for introducing me to the wonder world of
palms.
I would like to thank to Dr. Harry Wiriadinata, Titik Rugayah, Ir. Himmah
Rustiami, M.Sc. and Teguh Triono Sp. M,Sc., for helping me in various ways.
viii
'1 would like to thank Kartika Indah, S.Pd, M.Si, Dra. Kasrina, If. Nurhaida
Iriani Sinaga, Medi Hendra, S.Si., Andam Surianty Ardan, S.Si., for their moral
supports.
My very special thanks are due to my uncle and aunt (Drs. H. Arifin Saleh
and Dra. Hj. Nurlina Arifin) for giving me continuous moral support and attention
during my study in Bogor, and also to my sister Dra. Riwayati, M.Si, for her
attention and advice.
IX
TABLE OF CONTENTS
ABSTRACT ................................................................................... .ii
CURICULUM VITAE ...................................................................... vi
ACKNOWLEDGEMENTS ................................................................. vii
TABLE OF CONTENTS ................................................................... .ix
LIST OF FIGURES .......................................................................... xi
INTRODUCTION .............................................................................. 1
METHOD OF STUDY .................................................................... 3
RESULT AND DISCUSSION .......................................................... 5
Section P hyllanthectus Furtado ................................................... 7
Key to species of the Genus Calamus (Palmae) section Phyllanrhectus
Furtado in Sumatra .............................................................. 8
Species Description ................................................................. .12
Calamus asperrimlls Blume ............................................. 12
Calamus axillaris Becc ................................................... 13
Calamus caesius Blume .................................................. .15
Calams laevigatus Mart .................................................. 18
Calamus manan Miq ...................................................... 20
Calamus oxleyanus Teysm. et Binn ................................... 22
Calamus palembanicus Becc ........................................... 25
Calamus tllmidus Furtado ............................................... 26
Calamus unifarius H. Wend!.. ......................................... 27
x
CONCLUSION ... ........... ' ............................................................ .34
REFERENCES ............................................. '" ........................... 35