Upaya Meningkatkan Keragaman Somaklonal Tanaman Penghasil Gaharu (Aquilaria malacensis Lamk) secara Kultur In Vitro.
(/4ran d@4tr6rrnl)s{!!k rf ,i
dF!trtr'6hin]gnebyEjrdbim
^g$radlatttutb
Nt@4r L,bu d
*k
done
50n4nL\ ) !m@(e), 5!0pr([) ]000DniA4rj@;;h
r*^LBLoabis*o(5)h€Lsrtrroi.0,0op4NAA+o,o0p4lAp(Ko).
J,(pnNAA+5.00FrAp(rrI5,@ ppnNAA+10,00pN uF(re| j.oo
Fn NAA+15,000D BdlK) 6 5,o
diFo&'gd4ai*.ndjddoar0gd
l]dd{tsldDhl4nqiro[dsnat
$r!i he Fig Dhi ekoiomioFjuh
c4r!fudddih!uWDldbpuigjl@M.ydek@led
sejuAquhd4Fne'qdnidd&egi
ddfusFi6/qularuyhgdduifunPTdlrsi9h!uli6tfu'l6i
sb$a,, db tue, s*rylu a vdrsir ld:hh FDellir $ie dei Nsa
rssEreddh(ddshiBd(se M, re%) pohonGrun dis!ft@
d. s.hbA h$m4 sr!!o& Lina
FdhnididbdeP5i,4DFbstrD!L.nF'n'srisnfuiqsrlrru
bruo.mheded'Fdlgnghi$bea
iid!$iFrft n,bdikdupe@leo di ebebi
akue &! Bs n{rd?r dio rdu! [
ckF
srr@ rdmAa ) hss! r (sdo lc sbm kuLds $Fr ddsFi
iuPuh ddr *sportr (suDm, 2m2)
5
i!6
ellrudi|enod@h$fr$!neD]ilrd,
d]dbk.isdhhighqabmnm'
Flnsgfull(aibuoeaid!buirFnu
tur $drho bqb crubni jds,e/2,4
n*a d sru @jsisi d6h 06 (coDddM
Endried sFci{) Fd rqwE y s
FfuFd.n.*FnFbrodibr6
Ncac dnm rutu jenh h mr d
otr
rrtdiod rB& d
dienukei (turd{,Amr .Mdlrb da
DdltEebFhsdoFbagbn$b.e!i3uddAfu!e
$klisulkmdqb6q!d*5i4h$i!
Mreaijuts! dF lkri
bgfud{gibiliebloobtritb
6 umboh@F dn Araidi, 2m0
I rcbu erDro
ndgia boburdd l2rFbDFigb
s€rsjlds sade 20q de $hn,
e$
Mum
Fajar, Perm
dai
eru mb siruju4,
oeiunnkM ruurrr Fhor
Psrn
4h
ndruib!j,&ohcanqddif hntry! Dlyr k4snbihhtipureoh4,he\
y! ! $hbr 17 % (sou,re60; sari.. 20Gl sdosh, pde$ku hbi nd! ui
L4hn*sbdreF'']
jdi@
ldn
bibil
b'4e os'trd turidis@
e'rdjan,@ 6Fr diliikd $rd stu
' rEh!
do jffituh y@s ohr ltr4sl. $ngm
fhrir,ler4 drrln
dFlndi4btDk.dgllo$'n*hi
yog diF443s siiis edn! h€@
ttbMiryrfufue},ardd'hFJb'
sfuly!@ldFnigkadknsnugo,(roml
Fdiu oisisl hD jds lq-r{a
m6ddshbyqsm.ni]bF€Di
lduk PrcFm
besliiknogrEitkisubdstubeRsri$iqdjujn, he*qn&u
di
'u]dewberucfublFsnhi.kPi4s&igkldis!ruttl&El3@bd
ehdm Ebdf c!h4 dibidieb e4rd Fng di bq i4pi Fdi.in ra6r
reanai
@gisi
Fh.ru
tsoi4kxhk@mesondocl&!m
oe@luhr
bi,i
'dipr
k€I!hmo@gj&eabqhnF&ibcddunfi7.Ibhtr.t&4ra
Dur.
yhg
jld
€rLsl'dbuangt*'n@lhh.&tr!d,
ke
enh Fb4 di$h
! hhk n{ebd*h
b@y*,tsnmbdu!frn.id4grgm
brbi d! a n iudrrh
sisknii.&nFmbtnfury!hqagbbry$%
.kDk
hgkd sd
ndunL
bFde keBhki
F
hM nd^@ oenFrh
uFyamsicklrengfun$nak]o'frdduiFhkui2,4.D&oru.kfus
*'hn,ndazlegfuMbd,d]ii*U@ohbd
B'hm rlDi Gkpre) eq dlnbi dl
E
!.d.r,
dojuhF
bia yas
ditdruru
jad^F ddd l€lhbei
Ftunblbtr tu FL@b@ ekde n
bb]hlieid]siohoiimkngdot
(redh0, Ghp, 4.bih koNnd sidhii rebib rhBi
nd@!s *shn run&{rk r!6 Ghond)
&n
^uksh
ilk *o
b,kl,ufukmfr'ryb'kail.hgnol$n
shirs u
hF
eF dijdilai eh s
l!runli Fqb{ I$ho (!q/r4 n h4rr rrnk) eqn rL
rutsi F@1..i i dbb uduk: j)
fi
viro
AFt']rk@5q6iAuksi,.1D@s
dtlbku!fu3kliF&ndawrMFngd
dam
tr8k
DeLebnh
bffiro
roska
(FJ$6o Drea rutur Em &nh,
lkoionilj4giahhdalddblfro
h$n wlk
ndLojaos prcdlr1,i6
n@l,ldu FoFn
Fndod tmfr
P.Mktr ddni &n
d@)
)
$d. dimma
b
Ad hsrap GAJ 6@ 6 di e4iD. dina@ sri4 sd nc@crn nda
oh6!PDdlks]kbsPdlidiMM
htFifu :htrFzpr:5r2,4o:
r0 @2,4D:Dppm2,4D:r0@mrJ_D,
sri4$ diEobi
dhRBt&40sd'k64qd$hgi
yus e,nd, iFh dhij!*d lii BM F4 bd nve r %
n
&n
ro
P y!09
plr
r3.D + a
M
BAp
6
hoblkdtu.
rd
r,0 rlm 11 D +
d
5
hrltFrtu r!tsr!.D
ppm BAp,
'0,0
G@.
dm
h9
yqA
shhss
$hdra
be$& lFh dih',
tun. dn
5@F5m[@$@]rhpdgldfum.yh!nd'Pdi:
aI(er!)nrs
ds din !i I (dulmne
6$rdByfugnobg]d;
tu aErliki@ n{ullrko tsDsld etusikodsrd
BNr
r,4D
5%dbj s@darabrI
TellnemFnihful)olMFn6eckdosigidlPt.iw
dijuheri Fd Fnroo r p@ d.o r P
,$.00Dn,,opr,2J!r,fu10|@,'!D
tsdtu ar Fo4rr Nhbd x4 D Fe
ek$h sMru. Ab'di (ie4) m.@M
m*y&bhFn@@ttl0l0Fe
.lxeh ib *idid, jhr
6
rom$nt n
hlni*spbnUldiiqFirndhl
edir
d.hn
d!
{arwt c@ee &o strdoeloD. re&)
!44e hrdp dn Fk!6& .k
1 . D l.iiLtk;nr_G
rer rtu2
mhFrlM&! jue &
dsruk bus
rd!@
fujiiddugblhq&pliiFigdbidPdherdbr}sg
E'irl
dijumPr,
Fd!
\ho0i,
cros ds sheir,id tiq4) I do srrh (res) anFkkd
tonnld dksh 12,4-D)
hahH
p@aob!M[roE{r*i ,,4.D plnhorukrm kru t4d, i,b
rEir4dsd,
k,40dq
d
h
ddan
lrdd'Mo, l93I ] dn w]diAleIy.
s.rr^jurF
Fdeo
l)lhri *eho
ki&
$in&ng (Rio,sii,
a$)
I
rienbq|ulHs,*bgimmFdall
hiq knlneM dqh! h.n& d idr d
hju
ru$
lsys d4d
Mredi {1e:) Fns nmi rdapd F
ds rdh &€tu jdaen.krho bDui Fdi
LJ||fuhfuPda@fu|kh'do|goleBplqrn4b4|kkl^
sci sid* *cim osojurki Fogd berbqd ku{bsi
2,4D
e{ Fisd
drah dili &4d,
d stck+rdnen&dlkkr6rqEdijhF,F&
Fnaftoh,nLoenbi
DMNRTPa&'gdnFb5qod]$libFdaT.bl,
10.00 prm
l3
D,
di@'
hirbln:
doF*dh@*spJbm.nb(
iFh
dc
lJNFigsfl*hn]'gblfud0
cbe
&n scdhrotr oe&) | dli sdoa osr n MM Mtq
roE{rGi auki (2,4D) F3 1i@i E&. !o@r eFl neddus
FiMbfu beiuai ,J D
peob.{*Nr^
du rrjdi tib FidrFj
tr dhD ld{ siEbg (Rr0.s,4
kotlge, dl c[iisoi, l9e , &l
r*)
'
'9]di,sG'y
n@&dd l!16' *hFiroE Fndgl
Nd'neqdn cucwtr (191)
brus dh hju Fnuoruhlmy! @Fr
wfood,
bdbe eba rI!!' &n janoeo *$h
M^ydj (re2) Fig ie{ rdipd F
b!s lerhsidjdce elsdm rsBnu Fdi
44!s!!4!B!6
nrsr srhh rrii
tub kdiun be&&
iFh
lrrt
Hisi p4Fndm sddd d il
IujhE@ts]ljoopeDl'1D+5,(
N"...",-.
t,a
q4r
F6okrkh! n rciskk
\hhrod! (res) , &o sdrd (re9 o
l4einbryeDluFalF@trtnbd
($brd'@ h.eDFa i Jal @@ah $brd dlji &nen 3Nr
nFh 5 1; drajiki F( r rsbd I
F&d
'q&ikFdakombiEihtrtMis.o
5.00
rpn 14.D dn 5o0 PPn tuP
tiqt
; &^ stu tree)
oqglei rsrcor DentE d $@dc. hbih.iiei rFb a€rtdikBcinboi
wdrinru
srordh (!AP) bfuisl
n€n4sis *r (^ssr. res)
I1!!!
t4'ru
etan
uqtuu
44,s.
L
r{utlrlNok Fosr(rgs &re
rturro& kr" r(qd
(Perri
1!i hhp r)
Pn 13
rcdDe Fdr
Fd'io
bhp e&idsr kLu
D'r,0oDd !N
nouj 6i
t*rkdo
kL5 rEo6 eilkbG) ertrtu dlEoral
no@ukroDbianloi!Bo!J,000Fi:+D+5'@piBN
o 4n1r LrDoo FlDi i
ftm@Lw rq3 l.rn*kdb4dnp
$sn(e!d h,) F*!ja Ftuo& ued
Fo od.l?v 4
o
r{s! @!F:,4
D
'... ' 'rc
sdMi M re hisjftis r@bnu Fsb Fbn
;hBo
pEidin! dMn
dF!trtr'6hin]gnebyEjrdbim
^g$radlatttutb
Nt@4r L,bu d
*k
done
50n4nL\ ) !m@(e), 5!0pr([) ]000DniA4rj@;;h
r*^LBLoabis*o(5)h€Lsrtrroi.0,0op4NAA+o,o0p4lAp(Ko).
J,(pnNAA+5.00FrAp(rrI5,@ ppnNAA+10,00pN uF(re| j.oo
Fn NAA+15,000D BdlK) 6 5,o
diFo&'gd4ai*.ndjddoar0gd
l]dd{tsldDhl4nqiro[dsnat
$r!i he Fig Dhi ekoiomioFjuh
c4r!fudddih!uWDldbpuigjl@M.ydek@led
sejuAquhd4Fne'qdnidd&egi
ddfusFi6/qularuyhgdduifunPTdlrsi9h!uli6tfu'l6i
sb$a,, db tue, s*rylu a vdrsir ld:hh FDellir $ie dei Nsa
rssEreddh(ddshiBd(se M, re%) pohonGrun dis!ft@
d. s.hbA h$m4 sr!!o& Lina
FdhnididbdeP5i,4DFbstrD!L.nF'n'srisnfuiqsrlrru
bruo.mheded'Fdlgnghi$bea
iid!$iFrft n,bdikdupe@leo di ebebi
akue &! Bs n{rd?r dio rdu! [
ckF
srr@ rdmAa ) hss! r (sdo lc sbm kuLds $Fr ddsFi
iuPuh ddr *sportr (suDm, 2m2)
5
i!6
ellrudi|enod@h$fr$!neD]ilrd,
d]dbk.isdhhighqabmnm'
Flnsgfull(aibuoeaid!buirFnu
tur $drho bqb crubni jds,e/2,4
n*a d sru @jsisi d6h 06 (coDddM
Endried sFci{) Fd rqwE y s
FfuFd.n.*FnFbrodibr6
Ncac dnm rutu jenh h mr d
otr
rrtdiod rB& d
dienukei (turd{,Amr .Mdlrb da
DdltEebFhsdoFbagbn$b.e!i3uddAfu!e
$klisulkmdqb6q!d*5i4h$i!
Mreaijuts! dF lkri
bgfud{gibiliebloobtritb
6 umboh@F dn Araidi, 2m0
I rcbu erDro
ndgia boburdd l2rFbDFigb
s€rsjlds sade 20q de $hn,
e$
Mum
Fajar, Perm
dai
eru mb siruju4,
oeiunnkM ruurrr Fhor
Psrn
4h
ndruib!j,&ohcanqddif hntry! Dlyr k4snbihhtipureoh4,he\
y! ! $hbr 17 % (sou,re60; sari.. 20Gl sdosh, pde$ku hbi nd! ui
L4hn*sbdreF'']
jdi@
ldn
bibil
b'4e os'trd turidis@
e'rdjan,@ 6Fr diliikd $rd stu
' rEh!
do jffituh y@s ohr ltr4sl. $ngm
fhrir,ler4 drrln
dFlndi4btDk.dgllo$'n*hi
yog diF443s siiis edn! h€@
ttbMiryrfufue},ardd'hFJb'
sfuly!@ldFnigkadknsnugo,(roml
Fdiu oisisl hD jds lq-r{a
m6ddshbyqsm.ni]bF€Di
lduk PrcFm
besliiknogrEitkisubdstubeRsri$iqdjujn, he*qn&u
di
'u]dewberucfublFsnhi.kPi4s&igkldis!ruttl&El3@bd
ehdm Ebdf c!h4 dibidieb e4rd Fng di bq i4pi Fdi.in ra6r
reanai
@gisi
Fh.ru
tsoi4kxhk@mesondocl&!m
oe@luhr
bi,i
'dipr
k€I!hmo@gj&eabqhnF&ibcddunfi7.Ibhtr.t&4ra
Dur.
yhg
jld
€rLsl'dbuangt*'n@lhh.&tr!d,
ke
enh Fb4 di$h
! hhk n{ebd*h
b@y*,tsnmbdu!frn.id4grgm
brbi d! a n iudrrh
sisknii.&nFmbtnfury!hqagbbry$%
.kDk
hgkd sd
ndunL
bFde keBhki
F
hM nd^@ oenFrh
uFyamsicklrengfun$nak]o'frdduiFhkui2,4.D&oru.kfus
*'hn,ndazlegfuMbd,d]ii*U@ohbd
B'hm rlDi Gkpre) eq dlnbi dl
E
!.d.r,
dojuhF
bia yas
ditdruru
jad^F ddd l€lhbei
Ftunblbtr tu FL@b@ ekde n
bb]hlieid]siohoiimkngdot
(redh0, Ghp, 4.bih koNnd sidhii rebib rhBi
nd@!s *shn run&{rk r!6 Ghond)
&n
^uksh
ilk *o
b,kl,ufukmfr'ryb'kail.hgnol$n
shirs u
hF
eF dijdilai eh s
l!runli Fqb{ I$ho (!q/r4 n h4rr rrnk) eqn rL
rutsi F@1..i i dbb uduk: j)
fi
viro
AFt']rk@5q6iAuksi,.1D@s
dtlbku!fu3kliF&ndawrMFngd
dam
tr8k
DeLebnh
bffiro
roska
(FJ$6o Drea rutur Em &nh,
lkoionilj4giahhdalddblfro
h$n wlk
ndLojaos prcdlr1,i6
n@l,ldu FoFn
Fndod tmfr
P.Mktr ddni &n
d@)
)
$d. dimma
b
Ad hsrap GAJ 6@ 6 di e4iD. dina@ sri4 sd nc@crn nda
oh6!PDdlks]kbsPdlidiMM
htFifu :htrFzpr:5r2,4o:
r0 @2,4D:Dppm2,4D:r0@mrJ_D,
sri4$ diEobi
dhRBt&40sd'k64qd$hgi
yus e,nd, iFh dhij!*d lii BM F4 bd nve r %
n
&n
ro
P y!09
plr
r3.D + a
M
BAp
6
hoblkdtu.
rd
r,0 rlm 11 D +
d
5
hrltFrtu r!tsr!.D
ppm BAp,
'0,0
G@.
dm
h9
yqA
shhss
$hdra
be$& lFh dih',
tun. dn
5@F5m[@$@]rhpdgldfum.yh!nd'Pdi:
aI(er!)nrs
ds din !i I (dulmne
6$rdByfugnobg]d;
tu aErliki@ n{ullrko tsDsld etusikodsrd
BNr
r,4D
5%dbj s@darabrI
TellnemFnihful)olMFn6eckdosigidlPt.iw
dijuheri Fd Fnroo r p@ d.o r P
,$.00Dn,,opr,2J!r,fu10|@,'!D
tsdtu ar Fo4rr Nhbd x4 D Fe
ek$h sMru. Ab'di (ie4) m.@M
m*y&bhFn@@ttl0l0Fe
.lxeh ib *idid, jhr
6
rom$nt n
hlni*spbnUldiiqFirndhl
edir
d.hn
d!
{arwt c@ee &o strdoeloD. re&)
!44e hrdp dn Fk!6& .k
1 . D l.iiLtk;nr_G
rer rtu2
mhFrlM&! jue &
dsruk bus
rd!@
fujiiddugblhq&pliiFigdbidPdherdbr}sg
E'irl
dijumPr,
Fd!
\ho0i,
cros ds sheir,id tiq4) I do srrh (res) anFkkd
tonnld dksh 12,4-D)
hahH
p@aob!M[roE{r*i ,,4.D plnhorukrm kru t4d, i,b
rEir4dsd,
k,40dq
d
h
ddan
lrdd'Mo, l93I ] dn w]diAleIy.
s.rr^jurF
Fdeo
l)lhri *eho
ki&
$in&ng (Rio,sii,
a$)
I
rienbq|ulHs,*bgimmFdall
hiq knlneM dqh! h.n& d idr d
hju
ru$
lsys d4d
Mredi {1e:) Fns nmi rdapd F
ds rdh &€tu jdaen.krho bDui Fdi
LJ||fuhfuPda@fu|kh'do|goleBplqrn4b4|kkl^
sci sid* *cim osojurki Fogd berbqd ku{bsi
2,4D
e{ Fisd
drah dili &4d,
d stck+rdnen&dlkkr6rqEdijhF,F&
Fnaftoh,nLoenbi
DMNRTPa&'gdnFb5qod]$libFdaT.bl,
10.00 prm
l3
D,
di@'
hirbln:
doF*dh@*spJbm.nb(
iFh
dc
lJNFigsfl*hn]'gblfud0
cbe
&n scdhrotr oe&) | dli sdoa osr n MM Mtq
roE{rGi auki (2,4D) F3 1i@i E&. !o@r eFl neddus
FiMbfu beiuai ,J D
peob.{*Nr^
du rrjdi tib FidrFj
tr dhD ld{ siEbg (Rr0.s,4
kotlge, dl c[iisoi, l9e , &l
r*)
'
'9]di,sG'y
n@&dd l!16' *hFiroE Fndgl
Nd'neqdn cucwtr (191)
brus dh hju Fnuoruhlmy! @Fr
wfood,
bdbe eba rI!!' &n janoeo *$h
M^ydj (re2) Fig ie{ rdipd F
b!s lerhsidjdce elsdm rsBnu Fdi
44!s!!4!B!6
nrsr srhh rrii
tub kdiun be&&
iFh
lrrt
Hisi p4Fndm sddd d il
IujhE@ts]ljoopeDl'1D+5,(
N"...",-.
t,a
q4r
F6okrkh! n rciskk
\hhrod! (res) , &o sdrd (re9 o
l4einbryeDluFalF@trtnbd
($brd'@ h.eDFa i Jal @@ah $brd dlji &nen 3Nr
nFh 5 1; drajiki F( r rsbd I
F&d
'q&ikFdakombiEihtrtMis.o
5.00
rpn 14.D dn 5o0 PPn tuP
tiqt
; &^ stu tree)
oqglei rsrcor DentE d $@dc. hbih.iiei rFb a€rtdikBcinboi
wdrinru
srordh (!AP) bfuisl
n€n4sis *r (^ssr. res)
I1!!!
t4'ru
etan
uqtuu
44,s.
L
r{utlrlNok Fosr(rgs &re
rturro& kr" r(qd
(Perri
1!i hhp r)
Pn 13
rcdDe Fdr
Fd'io
bhp e&idsr kLu
D'r,0oDd !N
nouj 6i
t*rkdo
kL5 rEo6 eilkbG) ertrtu dlEoral
no@ukroDbianloi!Bo!J,000Fi:+D+5'@piBN
o 4n1r LrDoo FlDi i
ftm@Lw rq3 l.rn*kdb4dnp
$sn(e!d h,) F*!ja Ftuo& ued
Fo od.l?v 4
o
r{s! @!F:,4
D
'... ' 'rc
sdMi M re hisjftis r@bnu Fsb Fbn
;hBo
pEidin! dMn