Materials Directory UMM :Data Elmu:jurnal:L:Livestock Production Science:Vol65.Issue1-2.Jul2000:

82 E . Gerber Olsson et al. Livestock Production Science 65 2000 81 –89 for soundness, conformation, gaits under rider, free genetic trend in the population based on BLUP- jumping and jumping under rider. The stallions were Animal model evaluations is estimated with infor- also scored for temperament and general appearance mation from the RHQT and the trend has been ´ in the tests for gaits under rider, free jumping and increasing rapidly during the last 10 years Arnason jumping under rider. The stallions usually had to be et al., 1997. When considering horses born from approved at two out of three tests to be licensed for 1988 to 1992, the annual genetic progress for gaits breeding. Annually about 200 stallions were pre- and jumping, respectively, have been about 0.05 and inspected for conformation and soundness. About 0.03 genetic standard deviations per year. 40–60 were selected to take part in the stallion The purpose of the SPT is to select sires of the performance tests and on average 8–12 of these coming generation of successful sport horses. As stallions were finally approved for breeding. Ohlsson and Philipsson 1992 have concluded, In 1997 some changes were introduced in the performance at RHQT is highly correlated to later stallion performance test. The test is now done as a competition performance, it is of great interest to 9-day-long station test at 4 years of age at Flyinge. find out to what extent the SPT and RHQT results The stallions are preselected in the autumn at 3.5 are genetically related. years of age, for conformation, soundness and free The objectives of this study were to estimate jumping. The latter trait is therefore excluded from heritabilities, genetic and phenotypic correlations and the regular stallion performance test. The stallions repeatabilities for traits scored in the stallion per- are also judged by foreign test riders in the new test. formance tests as well as correlations between The SPT is used for phenotypic selection among stallion performance test results and 4-year-old field prospective breeding stallions, but until now it has performance records of their offspring. These param- not been investigated if heritabilities and variances eters are necessary for construction of integrated for the traits tested are high enough for an efficient indexes to be used for performance evaluations and selection. Because the SPT was done as repeated continued evaluations of stallions based on early tests it was important to investigate the re- progeny results. The parameters are also useful for peatabilities of the results, and thereby determine the determining the opportunities of improving or sim- effects of reducing the number of tests. The correla- plifying the stallion performance tests. tions between traits in the Swedish stallion per- formance test have not been estimated before and it would be interesting to know their interrelationships

2. Materials

in order to make optimal use of the information for selection purposes, which may indicate if the tests 2.1. Stallion performance test data can be further simplified and less costly. Riding horse quality tests RHQT, a 1-day field A dataset comprising 378 stallions evaluated at the test for 4-year-old horses, have taken place in SPT during 1979–1993 was analysed. The stallions Sweden since 1973. Annually about one third, or took part in one to three performance tests. The 500–1000, of all the 4-year-old riding horses in dataset contained observations from a total of 683 Sweden participate at the tests which take place at tests and was used to estimate heritabilities, genetic about 20 different places every autumn. The traits and phenotypic correlations and repeatabilities be- scored are health status, conformation, gaits under tween tests. The number of observations in the rider and jumping. A temperament and general dataset is shown in Table 1. All traits were given appearance score is given at the latter two parts of scores between 1 and 10. The temperament and the tests. All traits are given scores between 1 and general appearance scores for the two jumping traits 10. The tests are more precisely described by Gerber have only been recorded since autumn 1985 and the et al. 1997. The RHQT system aims at genetic corresponding score for gaits under rider since evaluations of stallions and mares and of providing autumn 1988. Also within a test some stallions were information about the quality of the individual young not shown in jumping under rider if the free jumping horses as a guideline for choice of sport horses. The result was poor. The scores for conformation and E . Gerber Olsson et al. Livestock Production Science 65 2000 81 –89 83 Table 1 Number of observations per trait and combining of traits in the data from the stallion performance test Trait 1 2 3 4 5 6 1 Walk, trot, gallop 536 a 2 Temp. gaits 217 217 3 Free jumping 458 596 4 Temp. free jumping 189 478 478 5 Jumping under rider 315 278 235 315 6 Temp. jumping under rider 174 239 236 264 266 a Temp., temperament and general appearance. soundness were not included in the material as the tions from the SPT contained 683 records for gaits stallions were pre-selected mainly on these criteria. and jumping. There were 234 stallions with offspring Means and standard deviations for the different traits in the RHQT. Of these, 136 had a minimum of five scored in SPT are given in Table 2. offspring in the test and were included in the The evaluated stallions had 160 sires and 167 correlation analysis with RHQT results. There were maternal grandsires, 60 of them appeared as both 2964 mares tested in the RHQT with offspring, that sires and maternal grandsires. Among the sires and also had participated in the RHQT. maternal grandsires 56 had themselves participated For estimation of the genetic correlations between in the SPT. SPT and RHQT traits new combined traits were created from the original ones. The trait ‘gaits’ was 2.2. Stallion performance test and riding horse created as the average of the four scores for walk, quality test data trot, gallop and temperament and general appearance, for the data from the RHQT and SPT. The score for To estimate the genetic correlations between traits ‘jumping’ was in the RHQT defined as the average tested at SPT and RHQT, respectively, the material of either two or four scores for free jumping and from the SPT, described above, and results from the jumping under rider both were not always re- RHQT during the period 1983–1993 were merged. corded, and temperament and general appearance Results for mares with foal at foot when tested were for the two disciplines whenever recorded. Approxi- excluded from the analysis. The final dataset for this mately 50 of the horses were tested for free analysis comprised 6674 horses tested for gaits under jumping only, 40 for jumping under rider only, and rider and for jumping in the RHQT. In the SPT, 327 the remaining had scores for both traits. In the SPT and 359 stallions were tested for gaits under rider data the average of all four scores for the jumping and jumping, respectively, and another 8559 horses traits were used. However, at the age of 4.5 years occurred in the pedigrees. The dataset with observa- and above, the free jumping test was exchanged for Table 2 Means, standard deviations S.D., minimum Min and maximum values Max and coefficients of variation CV of the different traits in the data from the stallion performance test Trait Means S.D. Min Max CV Walk 6.91 1.18 3 10 17.1 Trot 6.72 1.29 4 10 19.2 Gallop 7.09 1.09 5 10 15.4 a Temp. gaits 6.52 1.21 1 10 18.6 Free jumping 6.49 1.72 1 10 26.5 Temp. free jumping 6.56 1.87 1 10 28.5 Jumping under rider 6.70 1.56 2 10 23.3 Temp. jumping under rider 6.61 1.68 1 10 25.4 a Temp., temperament and general appearance. 84 E . Gerber Olsson et al. Livestock Production Science 65 2000 81 –89 Table 3 Number of horses N , means, standard deviations S.D., minimum Min and maximum Max values and coefficients of variation CV for the average scores of different traits in the combined data set from the stallion performance test SPT and riding horse quality test RHQT Trait average score N Mean S.D. Min Max CV Gaits RHQT 6674 6.52 0.89 3.00 9.33 13.7 Jumping RHQT 6674 6.79 1.50 0.50 10.00 22.1 Conformation RHQT 6674 7.47 0.49 5.00 9.60 6.6 Gaits SPT 136 6.83 0.92 4.33 9.67 13.5 Jumping SPT 136 6.24 1.63 1.00 10.00 26.2 another jumping test under rider. All tests at the the random effect of the kth tested stallion, mean50 2 different ages were included and means were calcu- and variance5 As ; pe is the random effect of the a kl lated for each created trait. The trait ‘conformation’ permanent environment, mean50 and variance5 2 in the RHQT data was defined as the average of five s ; e is the random residual effect, mean50 and pe ijkl 2 scores comprising type, head-neck-body, legs, walk variance5s . e and trot at hand. Table 3 contains means and Repeatability for the traits scored in the SPT was 2 2 2 2 2 standard deviations for the traits scored. estimated as s 1 s s where s 5 s 1 a pe P P a 2 2 2 s 1 s , and s is the additive genetic variance. pe e a Standard errors of the variance and covariance components were provided by the DMU programme

3. Methods