Isachne pulchella Roth ex Roem. Syst. Veg. 2: 476. 1817. Isachne pangerangensis Zoll. Mor. Syst. Verz. 1842—1844. 102. 1845.

Plants annual, mat forming, or caespitose. Culms up to 10 cm long, less than 1 mm thick, hollow, branches, rooting at the base. Nodes glabrous; internodes up to 2 cm, glabrous. Sheaths pilose. Ligule c. 0.9 mm long, a fringe of hairs. Blades 1.5 cm long by 5 mm long, lanceolate-ovate, both surface hispid, densely or sometimes inconspicuously appressed hairy, base rounded, apex acute. Inflorescence c. 4 – 14 cm long, a panicle with c. 14 spikelets, eglandular, contracted, elliptic, branches smooth. Spikelets c. 1.4 mm long, solitary, subglobose, green; pedicels c. 5 mm long; fertile spikelets pedicelled, comprising 2 fertile florets, without rachilla extension, orbicular, dorsally compressed, breaking up at maturity, disarticulating below each fertile floret. Glumes similar, deciduous, shorter than spikelet, thinner than fertile lemma. Lower glume 1.4 mm long, as long as upper glume, as long as spikelets, membranous, without keels, surface glabrous, or hispidulous, apex obtuse. Upper glume, as long as adjacent fertile lemma, membraneous, keels absent, the surface glabrous, or hispidulous, apex obtuse. Floret with fertile lemma elliptic, or orbicular, c. 1.2 mm long, coriaceous, without keels; lemma surface puberulous, margins involute, apex obtuse; palea coriaceous, 2-nerved. Anthers 3. Notes Found in China, Eastern Asia, Indo-China, and Malesia. Distribution at Sulawesi Central: Mt. Roroka Timbu-West slope, Alt. 1900 m, 0 30’ – 1 30’ S 119 30’ – 120 30’ E; Wera waterfall near Parampadende, c. 25 km SW of Palu, Alt. 250 m, 0 30’ – 1 30’ S 119 30’ – 120 30’ E; Mt. Nokilalaki, Alt. 1700-2200 m, 01 13’ S 120 08’ E. South: Masamba, Kamboena-top, Alt. 2860; Celebes O, O.A.; Masamba, o-zyde v. Takalar, Alt. 2400-2000 m; Todjamboe, boven Palopo, c. 700 m.; Rante Lemo, Alt.1500 m. Habitat Tiny herb creeping in mud of small pool, montane forest of Fagaceae + Myrtaceae, forming a mat on rocks base of waterfall, open areas. Specimen examined Eyma 1373, 1508 BO; Balgooy 3156, 3614 BO; Meijer 9867 BO; Steenis 10352 BO; Kjellberg 3003 BO.

50.9. Isachne pulchella Roth ex Roem. Syst. Veg. 2: 476. 1817.

Isachne dispar Trin. Sp. Gram. 18:, pl. 86. 1827. Isachne heterantha Hayata. Icon. Pl. Formosan. 7: 56-57, f. 28. 1918. Isachne nodibarbata Hochst. ex Steud. Henrard. Blumea 33: 464. 1940; Reinwardtia 122: 159-179. 2004. Plants annual. Culms slender, creeping, branching, rooting at the nodes, up to 20 cm long. Nodes bearded. Sheaths up to 1 cm long, hairy on margins, with fine tubercle-based hairs. Ligule a fringe of hairs. Blades lanceolate, 1-4 cm by 3-5 mm, margin cartilaginous, undulate, ciliate, hairy at base, apex acute, the surface puberulous, hairy on both sides. Inflorescence a terminal panicle. Panicle open, 3-10 cm long; primary branches spreading, sparsely divided, glandular. Spikelets solitary: fertile spikelets pedicelled. Pedicels filiform, glandular. Fertile spikelets comprising 1 fertile florets, without rachilla extension; elliptic, 1.5 mm long, dorsally compressed, breaking up at maturity, disarticulating below each fertile floret. Glumes similar, deciduous, shorter than spikelet, thinner than fertile lemma. Lower glume elliptic, 1- 1.2 mm long, as long as upper glume, herbaceous, no keels, 5-nerved, apex acute. Upper glume elliptic, herbaceous, no keels,7-nerved, apex obtuse. Florets: basal sterile florets male, with palea; lemma elliptic, 1.5 mm long, 5-nerved, obtuse. Fertile lemma elliptic, 1 mm long, coriaceous, pallid, no keel; surface pubescent, margins involute, apex obtuse. Palea coriaceous, 2-nerved. Lodicules 2, cuneate, fleshy. Anthers 3, 0.4 mm long. Caryopsis with adherent pericarp. Notes Found in China, Eastern Asia, India, Indo-China, Malesia, and Australia. Distribution at Sulawesi Celebes. Habitat Damp places in the forest. Specimen examined Kurniawan 102 WALL.

50.10. Isachne pangerangensis Zoll. Mor. Syst. Verz. 1842—1844. 102. 1845.

Isachne firmula Buse. Pl. Jungh. 3: 39. 1854. Isachne monticola Buse. Pl. Jungh. 3: 39. 1854; Reinwardtia 122: 159-179. 2004. Isachne rhignon Steud Ohwi. Bot. Mag. Tokyo 55: 541. 1941. Isachne rigida Nees ex Miq. Fl. Ned. Ind. 3: 461. 1857. Plants perennial, forming cushion-like tufts or mats. Culms up to 28 cm long, 1 mm thick, slender, hollow, glabrous. Nodes glabrous; internodes c. 5 cm long, glabrous. Sheaths ciliate along outer margin. Ligule densely awn up to 3 mm long, a fringe of longish hairs or wanting. Blades c. 9 cm by 6 mm, lanceolate, with rounded or obtuse base and acute tip, with narrow, pale, thickened, scabrid or long-ciliate margin, scaberulous beneath or on both sides. Inflorescence up to 12 cm long, spike- like panicle; panicle 2 – 3 cm, branches subsolitary with few short pedicelled spikelets. Spikelets obovate, obtuse, slightly longer than 2 mm long and as broad at maturity, with pedicels c. 2 mm long. Glumes subequal, the lower in the closed spikelets slightly longer than the florets, in the mature spikelets usually as long as the florets, distinctly 5 – 7-nerved, appearing apiculate or notched by inrolling of the margins when dry; upper glume usually obtuse, 5-nerved; both glumes glabrous or hairy towards the tips; lemma subequal, the lower slightly longer, the upper distinctly pedicellate on the terete rhachilla. Caryopsis 1.3 mm long, yellowish brown. Notes Found in Malesia. According to Backer and Backhuizen 1968 and Jansen 1953, this species is variable in habit and size, and sometimes forming low cushions on mountain-tops. Distribution North: Tomohon, Alt. 700 m. Central: Taripa Poso. Habitat Marsh, volcanic area. Specimen examined Eyma 4046 BO, Koorders 19792 β BO.

50.11. Isachne stricta Elmer. Leafl. Philipp. Bot. 2: 463. 1908.