Diplazium barbatum Christensen Diplazium batuayauense Praptosuwiryo, sp. nov.

198 A.C. 319. --- SUMATRA: Dr. Cramer 40; W.J.J.O. de Wilde and B.E.E. de Wilde-Dyfjes 12793; O. Posthumus s.n. 27 July 1925; J.A. Lörzing 14730; J.A. Lörzing 5342a; J. v. Borssum W. 2767; J.J. Afriastini 613; J.A. Lörzing 6894. --- MALAY PENINSULA: G.H. Addison SFN 37159.

9. Diplazium barbatum Christensen

Diplazium barbatum C.Chr. in C.Chr. Holttum, Gard. Bull. S.S. 7: 272. pl. 59. 1934. --- TYPE: Borneo, Mt. Kinabalu, Holttum SFN 25386 Holotype, BM n.v.; isotype, US; K n.v., SING n.v.. Rhizome short, erect. Stipe ca. to 30 cm long, light brown throughout, densely scales toward base; scales light brown, linear lanceolate, 3-1.4 mm long, 0,25-1 mm broad, concolours, without thickening black strands, margin toothed, teeth distinctly forked. Lamina broad lanceolate, ca. to 65 cm long, 19 cm broad, pinnae 10 pairs; lower shortly stalked to 3 mm long, pinnae pinnate, ovate lanceolate, to about 12 cm long, 5.3 cm broad, free pinnulae 2-4 pairs below pinnatifid apex of pinnae; pinnulae sessile, almost at right angle to costa, oblong, margin entire, ends acute-rounded, slightly toothed; texture papyraceous, rachis and costa densely minute scales; veins free, pinnate in the pinnulae; veinlets to 9 pairs, once forked. Sori elongate from near costule continuing to acroscopic branch of veinlets covers 23-34 of their length, indusia pale brown, margin entire, persistent. DISTRIBUTION. Borneo. ECOLOGY. Terrestrial in mountain forest. Elevation 1400-2100 m. SPECIMENS EXAMINED. --- BORNEO: J. M.S. Clemens 32607; R.E. Holttum 25386; M. Kato, G. Murata Y.P.Mogea. NOTES. As notified by Christensen Holttum 1934, D. barbatum is somewhat resembling D. speciosum in general habit, colour and texture, but very distinct by its densely squamose stipe, rachis and costae.. 199

10. Diplazium batuayauense Praptosuwiryo, sp. nov.

TYPE: Borneo, Central Kalimantan, Mts. Muller, Batau Ayau, Above S. Talikot Puhung Kucan, ca. 450 m, 14 June 2004, T.Ng. Praptosuwiryo 1927 holotype, BO. Rhizoma breve erectum. Stipites gracilis, 19-5-23 cm longae, 3 mm crassa, squamis caducus brunneus, linear-lanceolatus, 4-10 mm longae, 0.5 mm latis, margine irregularis dentibus furcatis cum filum niger spissescens. Lamina pinnata, lanceolatis, 41 cm longis, 20 cm latis, pinnae 15-jugatae infra apice pinnatifidus; pinnae inferiora stipitae ad 3.5 mm longis, superiora adnatus, lanceolatae, 11.2 cm longis, 2 cm latis, basi inaequaliter truncatae ad cuneatae, e margine 13-12 costam lobatae; lobi 4-5.5 mm latae supra basim, truncatae, leviter crenati. Rachides non gemmiferae, glabrae. Venulae distinctus in superficiebus ambabus, liberis, pinnatus in uno lobo, venae principalis angulum 55º cum costa formantes, venulae 4-5-jugatae, simpliciter. Sori medial or close to margin of lobes, cover 13-12 of veinlets length 3 mm length. Indusiis brunneis, concolour, persistens, ineteger. Rhizome short, erect. Stipe slender, 19.5 -23 cm long, 3 mm thick, fallen scales; scales brown, lineary lanceolate, 4-10 mm long, 0.5 mm broad, margin toothed, teeth apart, with thickening dark brown strand irregularly, glandular cells present,. Lamina pinnate, lanceolate in outline, 41 cm long, 20 cm broad, pinnae 15 pairs below pinnatifid apex of lamina; lower pinnae stalked 3.5 mm long, upper pinnae adnate, one pair basal pinnae bending downward with cut away base; pinnae lanceolate, widest at 13 part from basal, 11..2 cm long, 2 cm broad, base unequally truncate on lower pinnae, cuneate at upper ones, apex acuminate, margin lobed to within 4 mm of costa or 13-12 way to costa; lobus widest at base, 4-5.5 mm wide above base, ends truncate, slightly crenate. Rachise not gemmiferous, glabrous. Texture thin. Veins distinct on both surface, free, pinnate in the lobus, main vein forming angle 55° to costa, veinlets 4-5 pairs, all simple, reaching margin. Sori medial or close to margin of lobes, cover 13-12 of veinlets length 3 mm length. Indusia brown, persistent, margin entire. 200 PARATYPE. BORNEO: Central Kalimantan, Mts. Muller, Above S. Talikot Puhung Kucan, track to Batu Ayau, 450 m, 14 June 2004, T.Ng. Praptosuwiryo 1927a, 1927b, 1927c, 1927d, 1927e. ANATOMY. Transverse section of stipe near lamina: Vascular bundle form interrupted U-shaped, with an angle 115º, angle slightly ridges, end almost simple. CHROMOSOMES. 2n = 164 Cytotype: T.Ng. Praptosuwiryo 1927c, BOHB, 205 Cytotype: T.Ng. Praptosuwiryo 1909, BOHB. DISTRIBUTION. Borneo ECOLOGY. Terrestrial in lower montane forest. It is Grows on humus rich soil in shady places. NOTES. This species closely related to D. sorzogonense. Diplazium batuayauense differs from D. sorzogonense in the following characters: Scales sharply toothed with thickening dark brown stand irregularly, fronds are much smaller, never reaching 1 m, lacks the fibrillose scales on stipe and rachis, pinnae lobed only to ½ way to costa, sori not impressed. D. batuayauense is also similar to D. malaccense. ETYMOLOGY. The specific epithet is from the locality where this species found, Batu Ayau, the mountain forest of Mts. Muller in Central Kalimantan, Borneo.

11. Diplazium beamanii M.G. Price