225 much larger; probably both D. Smithianum and D. insigne are local derivatives of
D. accedens or D. proliferum.
30. Diplazium kunstlerii Holttum
Diplazium Kunstlerii Holttum, Gard. Bull. S.S. 11: 88. f. 2. 1940. –
Athyrium kunstlerii Holtt. Holttum, Rev. Fl. Malaya 2: 564. f. 335. 1966. ---
TYPE: R.E. Holttum SFN 31194 holotype, US. Rhizome suberect, about 2 cm diam., scales on younger part. Stipe 7 mm
diam at base, 70 cm long, glabrescent, dark brown, nearly black and scales at base; scales rounded. Lamina bipinnate, about 107 cm long, 60 cm wide; pinnae
about 11 pairs; larger pinnae 52 cm long, about 16 cm wide, on stalk to 11 cm long, upper pinnae with shorter stalk; lowest pinnules to about 5 cm apart, largest
ones 9.2 cm long, 2.7 cm broad, on stalk to 2 mm long, base broadly cuneate to truncate, margin lobed to 23 way toward costa; lobes slightly oblique, 5-7 mm
wide, base dilated, margin slightly toothed, apex rounded; texture thicker than in allied species; rachis gemmiferous at upper portion; veins in each lobe 6-9 pairs,
oblique, mostly simple, distinct but hardly prominent on both surface. Sori at middle part of the veins or nearly close the costules, basal acroscopic veinlets
usually diplazioid; indusia narrow, hardly evident in mature sori, very dark brown. DISTRIBUTION. Malaya, Java.
ECOLOGY. Locally abundant near streams in shady forest in the foothills. Ca. 900-1000 m.
SPECIMENS EXAMINED --- MALAY PENINSULA: R.E. Holttum SFN 31194. --- JAVA: Winckel 1556B.
31. Diplazium laevipes C.Chr. in C.Chr. Holttum
Diplazium laevipes C.Chr. in C.Chr. Holttum, Gard. Bull. S.S. 7: 271.
Pl. 58. 1934. TYPE: Borneo, Mt. Kinabalu, H. 25259 Holotype, SING. Stipe 55-75 cm long, ca. 3-8 cm diam.near base, glabrescent, dark brown
and scales towards base. Scales on stipes rounded, to 3.5 mm diam., dark brown, margin entire, more thick toward central. Lamina subdeltoid, bipinnatifid-
226 bipinnate, pinnae numerous; pinnae shortly stalked 5-15 mm long, oblong
subtriangular-ovate outline, basal pinnae smallest, oblong lanceolate, base subcordate-subttruncate, subbasal pinnae 32-37 cm long, 8-19 cm wide, pinnulae
to 10 pairs below pinnatifid or deltoid lobed apex of pinnae, acroscopic basal basal pinnulae a little reduced; pinnulae almost at right angle to costa, lower
shortly stalked less than 1 mm long, oblong subtriangular, 10 cm long, 3.5 cm broad, base subtruncate, apex acuminate with slightly toothed, margin lobed 13-
23 way to costule; lobus oblong, slightly oblique, to 4 mm above base, acroscopic basal a little reduce, ends truncate, entire-slightly toothed; texture stiff , blackish
on upper surface and dark brown beneath when dry; rachis and costa deeply grooved and rather tomentose above, gemmiferous, gemmae bearing at adjacent
between upper part of rachis and costa; veins pinnate in each lobe, free, not so distinct on both surface, forming an angle about 65-80º to costule; veinlets simple,
to 9 pairs, occasionally once forked on one-two pair basal lobus, forming an angle about 25-35º to costulet. Sori elongate from basal veinlet cover ¼-23 their
length, diplazioid on basal acroscopic; indusia dark brown, fragile. DISTRIBUTION. Borneo.
ECOLOGY. Occurring on forest floor in deep shade near river or ravine at 900-1300 m.
SPECIMENS EXAMINED. --- BORNEO. Kalimantan Timur: M. Kato, M. Okamoto, E.B. Walujo B-11261, B-9596, B-10748, B-11486, B-9625, B-
7399; M. Kato, M. Okamoto, K. Ueda, D. Darnaedi E.B. Walujo B-8345; M. Kato, M. Okamoto, K. Ueda E.B. Walujo B-7399.
NOTES. As pointed out by Christensen Holttum 1934, D. laevipes is closely related to D. spiniferum. The two species are quite similar in size, colour
and division, but the step of D. spiniferum is prckly, the segments all entire, the sori far from the costule, and the frond coriaceous.
227
32. Diplazium latisquamatum Holttum
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