Diplazium riparium Holttum Biosystematic Study Of The Fern Genus Diplazium In West Malesia

254 single elements irregularly apart and projecting 1.1-2.1 µm. Exine: often visible through perine, smooth. DISTRIBUTION. Java. ECOLOGY. Diplazium profluens usually grows in moist humus-rich soil at shady places of small river banks. NOTES. D. profluens is closely related to D. latisquamatum and shared characters as follow, scales on stipes round or ovate, entire, and nearly black; sori basalis and occupying only lower half or less of the veins, margin of pinnulae lobed to within 2-3 of costa. It is different from D. latisquamatum on veinlets 7-9 pairs, middle acroscopic veinlets also diplazioid, margin of pinnulae lobed to within 2-3 mm of costae 78-89 way to costa. This species is also similar to D. kunstlerii, but the two species is differentiated on their pinnulae incision. Pinnulae D. kunstlerii is usually lobed 23 way to costa, while pinnulae of D. profluens lobed 78-89 way to costa.

51. Diplazium riparium Holttum

Diplazium riparium Holttum, Gard. Bull. S.S. 11: 97. f.5. 1940; Tagawa K. Iwatsuki., Fl. Thailand 3: 454. 1988. --- Athyrium riparium Holttum Holttum, Rev. Fl. Malaya 2: 554. f. 326. 1966. --- TYPE: Malay Peninsula, Selangor, Semenyih, H.L. Hume 8186 holotype, SING. Rhizome short, erect. Stipe 25-52 long, brown when dry, black and scales at base; scales 4-8.5 mm long, 0.5-1 mm wide, dark brown or nearly black, margin entire. Occasionally with glandular cells. Lamina pinnate, pinnae 2-3 pairs, terminal one like the others; argest pinnae oblong, 25-5 cm long, 6 cm wide, shortly stalked to 4 mm long or sessile, base cuneate, margin entire, suddenly narrowed near apex, apex caudate; texture chartaceous; rachis without buds; costa rounded beneath, glabrous, grooved on upper surface; veins at angle about 60º to costa, in small group, each group of 3 veins, middle vein forked 1-3 times, the outer of vein group uniting each other near margin of pinnae or 15 or less length from margin. Sori elongate along 2-4 to each vein-group, those on the outer veins of the group extending almost from the costa to margin, rest shorter, acroscopic 255 outer veins usually diplazioid; indusia not so thin, pale brown, fragile, margin entire. ANATOMY. Transverse section of stipe near lamina: Vascular bundle continued V-shaped with an angle 65º, ends simple, not forming a ridge. CHROMOSOMES. 2n = 82 Cytotype: T.Ng. Praptosuwiryo 2147, BO, 123 Cytotype: T.Ng. Praptosuwiryo 1847, BO. DISTRIBUTION. Java, Sumatra, Malay Peninsula, Borneo. ECOLOGY. Occurring on in jungle, rock soil, usually in wet places and often on stream banks at elevation 60– 1250 m sea level. USE. Dayak people use the young frond as vegetable after cooking. SPECIMENS EXAMINED. --- SUMATRA: T.Ng. Praptosuwiryo 2003a, 2003b, 2007, 2038, 2089, --- BORNEO. Kalimantan Timur: M. Kato, M. Okamoto, K. Ueda, D. Darnaedi E.B. Walujo B-8126; B-8235; T.Ng. Praptosuwiryo 1838, 1847, 1894, 1916, 1940, 1952a, 2108, 2123a, 2123c, 2145, 2147b, 2191c, 2193b, 2193c, 2200. --- BORNEO: Kostermans 10448; M. Kato, M. Okamoto, K. Ueda, D. Darnaedi E.B. Walujo B-8126, B-8235; M. Kato, M. Okamoto E.B. Walujo B-10070. MALAY PENINSULA. Perak: King 7894; L. Wray, 3590, 3650. Johor: R. E. Holttum S.F.N. 18301. Kelantan: Md Nur s.n. 14 February 1924. Melaka: H.N. Ridley 1589; H.N. Ridley s.n. Dec.1899. Trengganu: R.E. Holttum SFN 15323.

52. Diplazium silvaticum Bory Sw.