Diplazium dilatatum Bl Biosystematic Study Of The Fern Genus Diplazium In West Malesia

213 SPECIMENS EXAMINED. SUMATRA: T.Ng. Praptosuwiryo 2491. NOTES. Diplazium desisquamatum is in a glance similar to the small plants of D. polypodioides. However the dense scales throughout the stipes with ovate –lanceolate and entire margin will differentiate from D. polypodioides fastly. The stipes of D. polypodioides are densely covered by lineary lanceolate sharply toothed scales at base only. Moreover D. desisquamatum is differ from D. .polypodioides in characters combination as follow: larger pinnulae lobed to ¾ way to costa and veinlets 6 pair or less, while D. polypodiodes has larger pinnulae lobed more deep close to costules and veinlets to 11 pairs. ETYMOLOGY. The species epithet is from the Latin densus and squamatus meaning scales are dense in illustrating the densely scales throughout the stipes and rachis.

20. Diplazium dilatatum Bl

ume Diplazium dilatatum Blume, En. Pl. jav.: 194. 1828; C.Chr. Holttum, Gard. Bull. 7: 273. 1934. Backer Posth., Varenfl. Java: 132. 1939; Holttum, Gard. Bull. S.S. 11: 85. 1940. Rhizome stout, short, erect. Stipe to 80 cm or larger, 8 mm or more thick near base, dark green when living, black and very densely scales at base; scales narrowly linier, to 15 mm long, 1 mm wide, yellowish brown at middle, blackish brown and sharply toothed at margin. Lamina bipinnate-tripinnatifid, about 90 cm long, 70 cm wide, variable in size; pinnae oblong, narrowing toward acuminate apex, to 58 cm long, 70 cm wide, pinnules to about 16 pairs below deltoid lobed apex of pinna; basal pinnules usually a little reduce; larger pinnules stalked to 5 mm long, to 18 long, 4 cm broad, oblong subtriangular with attenuate apex, basal lobes a little reduce, base truncate-cordate on stalked ones, lobed ½ way to costa or a little more; lobes slightly subdeltoid-semiorbicular, apex rounded, margin subentire or serrate, 15 by 9 mm, commonly less; rachis glabrescent, costa with scattered narrow brown scales to 20 by 3 mm; veins in lobes pinnate with 5-7 pairs of simple or forked veinlets. Sori elongate along veinlets from near base to 23 length 214 SPORE. Monolete, bilaterally symatrical made asymmetric by prine, heteropolar, polar outline excluding perine elliptical, sides convex; equatorial longitudinal view excluding perine concave-convex; equatorial transverse view, proximal face concave, distal face convex; perinate. E: 31.8545.6949.56±4.78; P: 19.8428.5534.62±3.89. Laesura: concealed by perine. Perine: alate-to costate-alate, irregular envelope separated from exine surrounds the spore; wing- like muri projected c.6 -15 µm µm, terminating margin are often ciliate; surface of perine smooth. Exine: visible through perine, smooth under SEM. ANATOMY. Stomata: Polocytic, copolocytic and seppolocytic. CHROMOSOMES. 2n= 123 Cytotype: T.Ng. Praptosuwiryo 1073 DISTRIBUTION. India, Burma, S. China, Taiwan, Ryuku, S. Japan, Indochina, Malesia throughout to N. Australia. ECOLOGY. Terrestrial. On humus rich soil of mountain slopes primary forest or mixed forest at 100-1800 m sea level. VERNACULAR NAMES. Pakis layung Javanese, paku beunyeur Sundanese. USE. The young fronds can be eaten as vegetable after cooking. SPECIMENS EXAMINED. --- JAVA: Adelbert 142; Alston 12771; Dillewgn 706; Donk s.n., 603; Lefebu 113; Matthew 610; Mogea 2344, 2345; Mousset 50, 760; Popta 225; Posthumus 3584, 3768, 3939; Sapiin 2660, 2713, 2717; Zippelius 239. --- BORNEO: M. Kato, G. Murata Y.P. Mogea B-3873; R.E. Holttum SFN 25555; M.Kato, G.Murata Y.P. Mogea B-3738; M. Kato, M. Okamoto, E.B. Walujo B-10036. NOTES. Kato 1995 recognized two varieties D.dilatatum in Japan, viz. var. dilatatum and var. heterolepis. The first variety has scales on stipe base lanceolate, to 20 mm long, black at margin, while the second variety has scales on stipe broadly lanceolate, 10-15 mm long, 1-3 mm broad, hardly black at margin. 215

21. Diplazium dolichosorum Copel.