263 NOTES. This species is similar to D. sorzogonenense. As stated by
Holttum 1940 and Kato 1994, it differs from D. sorzogonense in the toothed scales, non-fibrillose scales on the rachise, and non-impressed sori.
56. Diplazium spiniferum Alderw.
Diplazium spiniferum Alderw., Mal. Ferns Suppl.: 265. 1917; D.
muricatum Copel. C. Chr., Ind.Suppl. 27. 1913; Athyrium muricatum Copel.,
Philipp. Journ, VII
c
, 61. Rhizome short, erect. Stipe 65 cm long, 8 m diam, near base, scales,
spinuous, pale brown or rather stramineous, shining, black at base; scales rounded, ca. to 4 mm long and broad, entire. Lamina bipinate, 95 cm long, 58 cm broad,
pinnated pinnae 5 pairs, pinnatifid pinnae 3 pairs, simple pinnae 8 pairs below pinnatifid apex of lamina; lower pinnae oblong lanceolate, to 32 cm long, 9 cm
broad, free pinnulae to 9 pairs, commonly 8 pairs; pinnulae almost at right angle, adnate-sessile, oblong, base broadly cuneate, apex acute-acuminate, margin
usually lobed less than 14 way to costule or almost entire; basal lobus the largest, to 3 mm wide; rachis veins free, forming an angle about 45º to costule, veinlets to
4 pairs, commonly 3 pairs, simple, forming an angle about 10-15° to main veins. Sori at middle veinlest covers ¼-13 of theirs length, basal acroscopic diplazioid;
indusia dark brown, narrow, margin torn, persistent. SPORES. Monolete, bilaterally symmetrical made asymmetric by
perine, heteropolar; polar outline excludng perine transversely elliptical, side convex; equatorial longitudinal view concave-convex to plano-convex; equatorial
transverse view, proximal face planar to convex, distal face hemispherical; perinate. E: 31.9639.5247.77±4.18, P: 14.8723.9229.21±4.14. Laesura:
concealed by perine. Perine: micro-costate, densely echinate under SEM, costae broken, densely micro reticulate, coralline; irregular envelope, separated from
exine, surrounds the spore in densely coralline with enchinae project 1.5-2.3 µm; lacunae are very small, irregular polygons less than 0.5-1.5 µm across; heavily
fenestrate throughout under SEM, giving coralline appearance. Exine: visible through perine.
264 ANATOMY. Transverse section of stipe near lamina: Vascular bundle
form an uninterrupted U-shaped, with an angle 90º, base flat inward and ridges outward, angles distinctly ridges, ends ridges outward and forming an angle 130º.
CHROMOSOMES. 2n = 82 Cytotype: T.Ng. Praptosuwiryo 1896c, BO.
DISTRIBUTION. Borneo. ECOLOGY. Growing on humus rich soil or limestones area of mountain
forest at altitude 100-1300 m sea level. SPECIMENS EXAMINED. --- BORNEO. East Kalimantan: M. Kato
H. Wiriadinata No. B 4868; M. Kato H. Wiriadinata No. B 5845; M. Kato H. Wiriadinata No. B 5847; M. Kato H. Wiriadinata No. B 6428; M. Kato
H. Wiriadinata No. B 4818; M. Kato, M. Okamoto, K. Ueda, D. Darnaedi E.B.Walujo No. B 8277; M. Kato, M. Okamoto E.B. Walujo B-10735; M.
Kato, M. Okamoto E.B. Walujo B-10848; M. Kato, M. Okamoto E.B. Walujo B-10800; Kunio Iwatsuki, M. Kato, Gen Murata Y.P. Mogea B-3342;
Kunio Iwatsuki, M. Kato, Gen Muarata Y.P. Mogea B-1078; M. Kato, M. Okamoto E.B. Walujo B- s.n. Kalimantan Selatan: M. Kato, Gen Murata
Y.P.Mogea B-3731; M. Kato, Gen Murata Y.P. Mogea B-3739; B-3891; T.Ng. Praptosuwiryo
1878, 1879, 1880, 1945, 1955, 1900, 1878, 1879, 1880.
NOTES. Kato 1994 stated that in the large leaves the pinnules are lobed more than ½ to costule.
57. Diplazium squarrasum K. Iwats. M. Kato
