264 ANATOMY. Transverse section of stipe near lamina: Vascular bundle
form an uninterrupted U-shaped, with an angle 90º, base flat inward and ridges outward, angles distinctly ridges, ends ridges outward and forming an angle 130º.
CHROMOSOMES. 2n = 82 Cytotype: T.Ng. Praptosuwiryo 1896c, BO.
DISTRIBUTION. Borneo. ECOLOGY. Growing on humus rich soil or limestones area of mountain
forest at altitude 100-1300 m sea level. SPECIMENS EXAMINED. --- BORNEO. East Kalimantan: M. Kato
H. Wiriadinata No. B 4868; M. Kato H. Wiriadinata No. B 5845; M. Kato H. Wiriadinata No. B 5847; M. Kato H. Wiriadinata No. B 6428; M. Kato
H. Wiriadinata No. B 4818; M. Kato, M. Okamoto, K. Ueda, D. Darnaedi E.B.Walujo No. B 8277; M. Kato, M. Okamoto E.B. Walujo B-10735; M.
Kato, M. Okamoto E.B. Walujo B-10848; M. Kato, M. Okamoto E.B. Walujo B-10800; Kunio Iwatsuki, M. Kato, Gen Murata Y.P. Mogea B-3342;
Kunio Iwatsuki, M. Kato, Gen Muarata Y.P. Mogea B-1078; M. Kato, M. Okamoto E.B. Walujo B- s.n. Kalimantan Selatan: M. Kato, Gen Murata
Y.P.Mogea B-3731; M. Kato, Gen Murata Y.P. Mogea B-3739; B-3891; T.Ng. Praptosuwiryo
1878, 1879, 1880, 1945, 1955, 1900, 1878, 1879, 1880.
NOTES. Kato 1994 stated that in the large leaves the pinnules are lobed more than ½ to costule.
57. Diplazium squarrasum K. Iwats. M. Kato
Diplazium squarrasum K. Iwats. M. Kato, Acta Phytotax. Geobot. 31
4-6: 179. 1980. Rhizome stout, short, erect, 1.8 c, thick when dry. Stipe 33.5-36.5 cm
long, 5 mm thick, deeply grooved above, dark brown, densely scales throughout; scales ovate-lanceolate, 2.5-12 mm long, 0.25 – 2 mm broad, concolour, light
brown, margin sharply toothed, without thickening black strands. Lamina simply pinnate, oblong, 72 cm long, 24 cm broad, lateral pinnae 17 pairs below deltoid
deeply lobed apex of lamina; lower pinnae shortly stalked to 1.5 mm long, upper ones adnate-sessile, 4.2-7.2 cm apart, patent at right angle to rachise, oblong,
265 one pair basal pinnae reduced to 7.7 cm long, 2.8 cm broad, larger ones 11.3-15.5
cm long, 2.7-3.3 cm broad, suddenty norrowed at 23 part of base, base shallowly auricle, lower truncate, upper broadly cuneate, base of upper pinnae broadly
cuneate, margin entire, or slightly waved, apex acuminate; texture thin, but firm subherbaceous, lower surface scatterly bearing stelate trichome; upper surface
glabrous; rachise densely scales, scales like those on upper part of stipes; costa raised below, densely minutely scales; veins anastomousing, veins group forming
angle 50-60° to costa, each vein group of 3 veins, middle veins forked 4-5 times again, outer veins uniting with the nearest of outer veins of other veins group ¼-
13 way of costa veins anastomousing 23-34 way to costa, forming areola more or less 1.5 mm wide, inner veins also uniting with nearest veins. Sori
elongate along veinlets, cover ¼-45 of their length, opening towards both acroscopic and basiscopic, sometimes diplazioid on basal acroscopic or on uniting
veinlets; indusia narrow, light brown, concolour, margin entire, persistent, firm, opening when mature.
DISTRIBUTION. Borneo. ECOLOGY. On humus-rich slopes near stream in evergreen forest, at ca.
1200 a.s.l. SPECIMENS EXAMINED. --- BORNEO: M. Kato H. Wiriadinata B-
4822, 5030.
58. Diplazium subalternisegmentum Praptosuwiryo, sp. nov.
