Diplazium hewittii Copel. C.Chr

221 CHROMOSOMES. 2n = 123 Cytotype: T.Ng. Praptosuwiryo 2341, BO DISTRIBUTION. Java. ECOLOGY. This species is growing on moist humus-rich soil of shady places in mountain forest. In the Mt. Halimun, D. halimunense was found growing among D. donianum and D. bantamense. Ca. 1300 m. SPECIMEN EXAMINED. --- JAVA: T.Ng. Praptosuwiryo 749, 2341. NOTES. D. halimunense may be closely related to D. bantamense, but the last two species appears to differ from the first species. Diplazium halimunese has irregularly sharp toothed scales with thickening black strands, while the margin scales of D. bantamense are minutely and regularly toothed; Pinnae of D. halimunense are cuneate at base with margin entire, while pinnae of D. bantamense are round at base with subentire or serrate at posterior portion. It also resembles D. donianum, but D. halimunense has erect rhizome, while D. donianum with creeping rhizome. The two species have similarities in irregular sharp toothed scales and ovate-lanceolate pinnae. Cytological observation of one individual of D. halimunense TNgP 2341b showed 2n = 123 triploid. Whereas most indivual of D. bantamense from Java are tetraploid and only some individual oktoploid. One collection number of D. donianum planted in Bogor Botanic Gardens from Sumatra is tetraploid See Chapter 5. ETYMOLOGY. The specific epithet is formed from name of the locality where this species is found, Mt. Halimun, Halimun National Park, West Java.

27. Diplazium hewittii Copel. C.Chr

Diplazium hewittii Copel. C.Chr., Index Fil. Suppl. 1: 26. 1913.; C.Chr. Holttum, Gard. Bull. S.S. 7: 273. 1934. Athyrium paripinnatum Copel., Philip. J. 11 2: 147. 1915. Diplazium paripinnatum Alderw., Handb. Suppl. 1 : 266. 1916. Rhizome short, erect. Scales on stipes concolour, dark brown-blackish, shiny, ovate, to 2.5 mm long, 2 mm broad, margin entire, glandular cell on tip when young. Stipe 25-67 cm long, 2.5-3.5 mm diam. near base, light brown, slightly muricate, black at base. Lamina pinnate – bipinnate, subtrangular in 222 ouline, 30 – 70 cm long, 20-27 cm broad. Pinnae opposite-subopposite; lower pinnae stalked to 1.5 cm long, lanceolate, 22 cm long, 5.7 cm broad, pinnate, apex acumimate; lower pinnulae adnate – subsessile, upper pinnule sessile below deeply lobed apex of pinnae; larger pinnulae nearly hastate, 1.8-3.3 cm long, 0.4 – 1.0 cm broad, base broadly cuneate, upper base distinctly auricle, lower base less, apex acute-acuminate, margin crenate; middle pinnae with pinnulae sessile; upper pinnae gradually less pinnate-pinnatifid; texture subcoriaceous; surface naked, lower pale, upper dark green when living; rhacise naked; veins free, forked in each crenatiron, oblique. Sori elongated along veinlet from near costule covers 56 way to margin, acroscopic branch diplazioid; indusia brown, fragile, persistent, margin entire. CHROMOSOMES. 2n = 123 Cytotype: T.Ng. Praptosuwiryo 1913b, BO. DISTRIBUTION. Borneo ECOLOGY. Growing on humus rich soil, low sandy ridges, on shaded places in the forest at 400-450 m. SPECIMENS EXAMINED. --- BORNEO: M. Kato, M. Okamoto, K. Ueda E.B. Walujo B-7383; M. Kato, M. Okamoto, K. Ueda E.B. Walojo B- 7902; B-7903. K. Ueda D. Darnaedi B-8835; T. Ng. Praptosuwiryo 1913a; 1913b; 1913c; 1908a; 1908b; 1908c; 1942; 2171; 2172a; 2172b, 2174; 2177; 2178; 2179; 2180. NOTES. Christensen Holttum 1935 stated that a series of specimens collected by Mjoberd in Sarawak has shown that Athyrium Hewittii, A. sarawakense , and A. paripinnatum of Copeland probably all forms of the same species, different in size but otherwise scarcely distinguishable. My collections from Muller Range, Borneo, showed that this species has variation in morphology from a young to adult. Young plants in which spores have not been produced are showing pinnate fronds while adult plants having fronds with pinnate-bipinnate. 223

28. Diplazium hottae Tagawa