Diplazium malaccense Presl Biosystematic Study Of The Fern Genus Diplazium In West Malesia

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36. Diplazium malaccense Presl

., Diplazium malaccense Presl, Epim.: 86. 1849; Tard. C.Chr. in Fl. Gén. I.-C. 72: 258. 1940; Tagawa K. Iwats., Southeast As. St. 5 : 104. 1967; Acta Phytotax. Geobot. 23 : 56. 1968. -- Athyrium malaccense Presl. Hottum, rev. Fl. Malaya 2 : 525. 1955. Rhizome short, ca. 1.5-1.8 cm diam, erect, densely scales on young part. Stipe 39- 48.5 cm long, 0.3-0.5 cm diam near base, densely scales at base; scales 10-12 mm long, 1 mm broad, lineary triangular, brown. Lamina pinnate, oblong elliptical –subtriangular in outline, 33-64.5 cm long, 15-30 cm broad, pinnae 16- 19 pairs below lobed apex of lamina. Lower pinnae stalked 2-5 mm long, 1-2 pairs basal pinnae bending downward, lineary- subtriangular, 9-19.5 cm long, 1.1- 2.8 cm broad, base very unequal, lower base cut away, upper truncate, upper pinnae with base subequally truncate or cuneate, Pinnae thin in texture, drying rather light green, margin lobed 13-23 way to the costa, apex acuminate; lobes about 4-7 mm wide at base, oblique, apex rounded to subtruncate, slightly toothed. Veins 4-7 pairs in each lobe, all simple. Sori from near base of veins to near margin of lamina, acroscopic basal sorus usually diplazioid. Indusia medium brown, thin but firm, narrow, margin entire. SPORES. Monolete, bilaterally summetrical made asymmetric by perine, heteropolar; polar outline excluding perine elliptical; equatorial longitudinal view planar convex to concave-convex; equatorial transverse view, proximal face concave, distal view convex. Laesura: concealed by wing-like muri. E: 37.7334.2427.53±3.03; P: 26.0120.8013.33±2.82. Perine: alate to costate-alate, often loosely reticulate; irregular envelope separated from exine surrounds the spore in continuous anastomosing wings, forming a loose reticulation; wing-like muri project 1.5-8 µm, terminating margins sparsely ciliate. Exine: visible though perine, smooth granulate under SEM. ECOLOGY. In evergreen forest, terrestrial on rather moderate slope, in light shade, on humus-rich soil ground. 20 – 1000 m. SPECIMENS EXAMINED. --- MALAY PENINSULA. Pahang: M.R. Henderson 11235; Md Nur 10508; R.E. Holttum 20788; A.B. Murdok s.n. 12 June 1913; H.N. Ridley s.n. 1891. Negeri Sembilan: R.E. Holttum 9565; Md 234 Nur s.n. 29 Nov. 1923. Melaka: A. Derry 14. Penang: R.E. Holttum SFN 31196. --- BORNEO. Kalimantan Timur: K. Ueda D. Darnaedi B-8982; M. Kato, M. Okamoto E.B. Walujo B-9070; M. Kato, M. Okamoto, K. Ueda E.B. Walujo B-7398; M. Kato, M . Okamoto E.B. Walujo B-10909; M. Kato, M. Okamoto, K. Ueda, E.B. Walujo B-7871; K. Iwatsuki, M. Kato, G. Murata Y.P. Mogea B-2472. Kalimantan Selatan: M. Kato, G. Murata Y.P. Mogea B- 4155; 4175. --- SUMATRA: W.J.J.O. de Wilde B.E.E. de Wilde – Duyfjes 12560; W.J.J.O. de Wilde B.E.E. de Wilde – Duyfjes 12920; T.Ng. Praptosuwiryo 2218, 2220, 2209, 2217, T.Ng. Praptosuwiryo 2223, 2224, 2225, 2226, 2229, 2230, 2233, 2235, 2236, 2237, 2238.

37. Diplazium megasegmentum Praptosuwiryo, sp. nov. Plate 7.