200 PARATYPE. BORNEO: Central Kalimantan, Mts. Muller, Above S.
Talikot Puhung Kucan, track to Batu Ayau, 450 m, 14 June 2004, T.Ng. Praptosuwiryo 1927a, 1927b, 1927c, 1927d, 1927e.
ANATOMY. Transverse section of stipe near lamina: Vascular bundle form interrupted U-shaped, with an angle 115º, angle slightly ridges, end almost
simple. CHROMOSOMES. 2n = 164 Cytotype: T.Ng. Praptosuwiryo 1927c,
BOHB, 205 Cytotype: T.Ng. Praptosuwiryo 1909, BOHB. DISTRIBUTION. Borneo
ECOLOGY. Terrestrial in lower montane forest. It is Grows on humus rich soil in shady places.
NOTES. This species closely related to D. sorzogonense. Diplazium batuayauense
differs from D. sorzogonense in the following characters: Scales sharply toothed with thickening dark brown stand irregularly, fronds are much
smaller, never reaching 1 m, lacks the fibrillose scales on stipe and rachis, pinnae lobed only to ½ way to costa, sori not impressed. D. batuayauense is also similar
to D. malaccense. ETYMOLOGY.
The specific epithet is from the locality where this species found, Batu Ayau, the mountain forest of Mts. Muller in Central
Kalimantan, Borneo.
11. Diplazium beamanii M.G. Price
Diplazium beamanii M.G. Price, Contr. Univ. Mich. Herb. 16: 193, f. 1-3.
1987. Type: Borneo: Pinosuk Plateau, 1400 m, Beaman 10724 Holotype, MICH; Isotype , K., MSC n.v..
Rhizome erect. Stipe stramineus, 66 cm long, 7 mm thick, densely scales at base; scales pale brown, ovate, 10 mm long, 4.5 mm broad, margin entire.
Lamina ovate, 60 cm long, bipinnate to deeply tripinnatifid at base, pinnae ? pairs; pinnae stalked to 2.5 cm long, subbasal the largest, to 37 cm long, ca. 17 cm
broad, pinnulae to 13 pairs, basal acroscopic slightly reduced; pinnulae stalked to ca. 1 mm long, 2 cm distance, subtriangular-lanceolate, to 8 cm long, 2 cm broad,
base truncate, apex acuminate, margin lobed to 56 way to costa within 2 mm of
201 costa; segments to ca. 5 mm wide, ends subtruncate, margin slightly denticulate-
crenulate; veins free, pinnate, veinlets to 5 pairs, usually simple. Sori from near costule cover 13 way of veinlets 1.0-2.5 mm long, basal acroscopic often
diplazioid; indusia broad, margin irregularly fringed, persistent. DISTRIBUTION. Borneo.
ECOLOGY. Terrestrial. Lower montane forest by stream. Elevation: 1400 m. Endemic to Mount Kinabalu.
SPECIMEN EXAMINED. --- BORNEO: Beaman 10724.
12. Diplazium betimusense Alderw
. Diplazium betimusense
Alderw., Bull. Buit. III. 2. 142. 1920. ---TYPE: J.A. Lörzing 5718 Holotype, BO; Isotype, L, Sumatra, Sibolangit, Betimus
River. Rhizome short, erect, 17 mm thick. Stipe dark brown, 48 cm long,
sparingly muricate, scales towards the base, scales on stipes dark brown, rounded- ovate, margin entire, deciduous, leaving the stipes roughish by their persistent
bases, 3.5 mm long, 3 mm broad. Lamina bipinnate, ? cm long, ? cm broad, pinnae ? pairs; lower pinnae stalked to 5.3 cm long, oblong subtringular, 40 cm
long, 18 cm broad, pinnulae 6-9 pairs below pinnatifid deltoid apex of pinnae; lower pinnulae shortly stalked to 1.5 mm long, upper ones adnate-sessile,
lanceolate, 8-10.5 cm long, 2-3 cm broad, base subequally truncate-cuneate,apex acuminate-caudate, sharply toothed, margin lobed to within 7 mm of costa or 13-
12 to cosule; lobus more or less widest at base, 5-6.5 mm broad, ends truncate, slightly toothed; rachise dark brow, glabrous; texture firm; veins free, pinnate in
the lobus, main veins forming angle 50-55° to costule, veinlets 4-5 pairs, forming angle about 10-15 º to main veins, distinct on both surface, simple, all reaching
the margin. Sori subbasal or medial, covers to 13 of their length, sometimes diplazioid on basal acroscopic veinlets; indusia narrow.
DISTRIBUTION. Sumatra. ECOLOGY. Growing on shade part of forest, near river at ca. 400 m sea
level.
202 SPECIMENS EXAMINED. --- SUMATRA: J.A. Lörzing 5718
13. Diplazium christii C.Chr.
