Upper glume lanceolate, 1.5-2.5 mm long, 1.5-2.5 times length of adjacent fertile lemma, membraneous, no keels, 1-nerved; primary nerves distinct; lateral nerves
absent; surface hispidulous, hairy in lines; apex acuminate, 1-awned 5-15 mm long. Florets: fertile lemma lanceolate, 0.5-1 mm long, hyaline, no keel, 1-nerved; apex
acute; palea hyaline, no nerved, no keels. Anthers 3. Caryopsis with adherent pericarp.
Notes
Found in China, India, Indo-China, and Papua.
Distribution at Sulawesi
Central: Palu, Pakuli, Alt. 50 m; Toboli, Alt. 600 m.
Habitat Specimen
examined
Posthumus 2255
BO; leg. ign. 48 BO.
93. ZOYSIA Willd. Type: Zoysia pungens Willd. Ges. Naturf. Freunde Berlin Neue
Schriften 3: 440-441. 1801. Brousemichea Bal. J. Bot.Morot 48: 163. 1890; Contr. U.S. Natl. Herb. 41:
240. 2001. Matrella Pers., Syn. Pl. 1: 73. 1805.
After Karl von Zois, 1756-1800, country gentleman and plant collector of Carniola, Austria.
About 10 species, Africa, Asia, Australasia, Pasific, North and South America. Plants perennial, rhizomatous, or stoloniferous, mat forming. Culms arising from
nodes along stolons, often densely branched at ground level. Ligule a ciliate membrane, or a fringe of hairs. Blades usually glabrous abaxially, adaxial surfaces
glabrous, scabrous, or sparsely pilose, apices often sharply pointed. Inflorescence composed of racemes. Racemes single, linear, or lanceolate, bearing few fertile
spikelets, or many spikelets. Spikelets packing crowded, or continuous; appressed, or ascending, solitary. Fertile spikelts pedicelled; comprising 1 fertile florets, without
rachilla extension; laterally compressed, 2-6 mm long, falling entire. Glumes dissimilar. Lodicules absent. Anthers 3.
Key to the Species
1.a. Culms up to 25 cm long; inflorescence up to 4 cm long; spikelets ascending; upper glume 3-nerved; palea present; anthers 1.5 mm long ..............................
.........................................................................................93.1. Zoysia matrella
b. Culms up to 10 cm long; inflorescence up to 1.5 cm long; spikelets loosely
overlapping; palea absent; anthers 0.7 mm long ..............93.2 Zoysia pasifica 93.1.
Zoysia matrella L. Merr. Philipp. J. Sci. 74: 20, 230. 1912
Agrostis matrella L. Mant. Pl. 2: 185. 1771; Blumea 261: 169-175. 1980; Grass. Burma, Ceylon, India Pakistan i-xviii, 1-767. 1960.
Zoysia malaccensis Gand. Bull. Soc. Bot. France 667: 303. 1919; Blumea 261: 169-175. 1980.
Plants perennial, mat forming, rhizomatous, and stoloniferous. Culms 5-25 cm long, wiry. Sheaths longer than adjacent culm internode; glabrous on surface. Ligule
a ciliate membrane. Blades 2-10 cm by 1-3 mm, filiform, or linear, flat, or involute. Inflorescence composed of racemes, single, 0.5-4 cm long. Spikelets ascending,
solitary. Fertile spikelets pedicelled; comprising 1 fertile florets, without rachilla extension; lanceolate to ovate, laterally compressed, 2-4 mm long, falling entire.
Glumes dissimilar, exceeding apex of florets, firmer than fertile lemma, shiny. Upper glume lanceolate to ovate, as long as spikelet, coriaceous, no keels, 3-nerved; apex
acute, muticuous; 1-awned, 1 mm long. Florets: fertile lemma oblong, 2-3 mm long, hyaline, no keels, 1-nerved; apex emarginate; palea 0.5 times length of lemma,
hyaline, no keels. Anthers 3, 1.2 mm long.
Notes
Found in Africa, China, Eastern Asia, India, Indo-China, Malesia, Australia, Pasific, North and South America.
Distribution
Central: Palu, Alt. 0 m, 50 m. South: Bonthain. Southeast: Kendari.
Habitat Specimen examined
Henippman 5023 BO; Posthumus 2256 BO; Kjellberg 3082, 382 BO.
93.2. Zoysia pasifica Goudswaard M. Hotta Kuroki. Acta Phytotax. Geobot.
